2026. 9.
금강산엉겅퀴로 알려져 있던 분류군이 반곡엉겅퀴로 새롭게 연구되었다.
한국식물분류학회지 56(3). 2026.
ABSTRACT: Cirsium reflexiphyllarium is described as a new species from Mt. Hwaaksan and Mt. Myeongjisan in Gapyeong-gun, Gyeonggi-do, and Mt. Daeamsan in Yanggu-gun, Gang won-do, Korea. This species has previously been misidentified as C. schantarense, C. dia mantiacum, and C. pendulum owing to its nodding capitula and occurrence in high-elevation habitats, and it also resembles C. pendulum in having nodding capitula. It differs from C. schantarense in having a non-arachnoid abaxial leaf surface, bracteate capitula, reflexed, non-glutinous phyllaries, and a linear pappus apex; from C. diamantiacum in the relative lengths of the phyllaries, corolla, and pappus; and from C. pendulum in its semi-amplexicaul leaf base and the ratio of the corolla tube length to the corolla limb length. Based on this combination of morphological characters, this species is recognized as a distinct new spe cies. The specific epithet reflexiphyllarium refers to the reflexed phyllaries. A description, illustrations, photographic plates, and an identification key are provided
요약: 경기도 가평군의 화악산과 명지산, 그리고 강원도 양구군의 대암산에서 발견된 신종 Cirsium reflexiphyllarium을 기재한다. 이 종은 이전에는 두상화가 아래로 숙여지는 형태와 고지대 서식지에서의 자생 때문에 도깨비엉겅퀴 C. schantarense , 금강산엉겅퀴 C. diamantiacum, 큰엉겅퀴 C. pendulum으로 오분류되었으며, 두상화가 아래로 숙여지는 형태는 C. pendulum과 유사하다. C. schantarense와는 잎 뒷면이 거미줄 모양이 아니고, 두상화에 포엽이 있으며, 포편이 뒤로 젖혀지고 점액이 없으며, 갓털 끝이 선형이라는 점에서 구분된다. C. diamantiacum과는 포엽, 꽃잎, 갓털의 상대적인 길이에서 구분된다. C. pendulum과 유사한 특징으로는 잎자루가 반쯤 감싸는 형태와 꽃부리통 길이와 꽃부리잎 길이의 비율이 있다. 이러한 형태학적 특징들의 조합을 바탕으로 이 종은 별개의 새로운 종으로 인정된다. 종명인 reflexiphyllarium은 뒤로 젖혀진 포엽을 의미한다.
분포지
도깨비엉겅퀴: 한국 분부지방 중국 연변과 러시아 극동
금강산엉겅퀴: 금강산 지역
반곡엉겅퀴: 강원도, 경기 북부 고산지역(화악산, 명지산, 대암산)
고개를 숙인 꽃차례를 가진 Cirsium reflexiphyllarium 및 관련 분류군의 식별 키
1. 줄기잎 밑부분은 줄기를 감싸지 않으며, 꽃부리통은 꽃부리잎 길이의 약 2.5배이다. ··················· C. pendulum 큰엉겅퀴
1. 줄기잎 밑부분은 반쯤 줄기를 감싸고 있으며, 꽃부리통은 꽃부리잎 길이의 2.5배 미만이다.
2. 잎 뒷면은 거미줄처럼 빽빽하게 덮여 있고, 두상화서는 포편이 없으며, 포엽은 밀착되어 있고 점액질이며,
갓털 끝은 곤봉형이다. ··············· ······························································ · C. schantarense 도깨비엉겅퀴
2. 잎 뒷면은 거미줄 모양이 아니며, 두상화서에는 포편이 있고, 포편은 뒤로 젖혀져 있으며 점액질이 없고, 갓털 끝은 선형이다.
3. 안쪽 포편은 바깥쪽 포편보다 길이가 두 배 길고, 꽃잎은 꽃부리통보다 길이가 약 두 배 길며, 갓털은 꽃부리통과 길이가
거의 같다. ························································································· C. diamantiacum 금강산엉겅퀴
3. 안쪽 포편과 바깥쪽 포편의 길이는 거의 같고, 꽃잎은 꽃부리통의 약 2.5배이며, 갓털은 꽃부리통의 약 3배이다.
····································································································· C. reflexiphyllarium 반곡엉겅퀴
| Key to Cirsium reflexiphyllarium and related taxa with nodding capitula 1. Cauline leaf bases non-amplexicaul; corolla tube approximately 2.5 times as long as the corolla limb ····· ········································· C. pendulum 큰엉겅퀴 1. Cauline leaf bases semi-amplexicaul; corolla tube less than 2.5 times as long as the corolla limb 2. Abaxial leaf surface densely arachnoid; capitula ebracteate; phyllaries adpressed, glutinous; pappus apex clavate ················· C. schantarense 도깨비엉겅퀴 2. Abaxial leaf surface non-arachnoid; capitula bracteate; phyllaries reflexed, non-glutinous; pappus apex linear 3. Inner phyllaries twice as long as outer phyllaries; corolla limb approximately 2 times as long as the corolla tube; pappus nearly equal in length to the corolla tube ······ C. diamantiacum 금강산엉겅퀴 3. Inner and outer phyllaries nearly equal in length; corolla limb approximately 2.5 times as long as the corolla tube; pappus approximately 3 times as long as the corolla tube ··· C. reflexiphyllarium 반곡엉겅퀴 |
도깨비엉겅퀴와는 경생엽이 난형-난상 피침형으로 잎 뒷면 맥 위에 짧은 털이 있으며, 두상화서에 포가 달리고, 총포편에 점성이 없으며, 뒤로 젖혀지는 점에서 구분된다.
금강산엉겅퀴와는 줄기에 조밀한 털이 있고, 잎 뒷면 맥 위에만 털이 있으며, 내포편과 외포편의 길이는 거의 같고, 관모와 화관 통부 갈이비가 3:1인 점에서 구분된다.
2010년에 만났던 반곡엉겅퀴
2022년에 다시 만난 반곡엉겅퀴
엉겅퀴(해안형) : https://qweenbee.tistory.com/8890179 https://qweenbee.tistory.com/8886916
https://qweenbee.tistory.com/8901298 https://qweenbee.tistory.com/8902508
https://qweenbee.tistory.com/8901337 https://qweenbee.tistory.com/8903881
엉겅퀴 : https://qweenbee.tistory.com/8901209 https://qweenbee.tistory.com/8901292
https://qweenbee.tistory.com/8901294 https://qweenbee.tistory.com/8902727
https://qweenbee.tistory.com/8903133 https://qweenbee.tistory.com/8903211
https://qweenbee.tistory.com/8888344 https://qweenbee.tistory.com/8904573
https://qweenbee.tistory.com/8905655 https://qweenbee.tistory.com/8906462
엉겅퀴 열매 : https://qweenbee.tistory.com/8897983
엉겅퀴 새싹 : https://qweenbee.tistory.com/8901216 https://qweenbee.tistory.com/8900675
https://qweenbee.tistory.com/8902169 https://qweenbee.tistory.com/8903617
https://qweenbee.tistory.com/8904902 https://qweenbee.tistory.com/8906799
개엉겅퀴 : https://qweenbee.tistory.com/8900255 http://blog.daum.net/qweenbee/8889405
https://qweenbee.tistory.com/8901291 https://qweenbee.tistory.com/8902727
https://qweenbee.tistory.com/8903480 https://qweenbee.tistory.com/8903514
https://qweenbee.tistory.com/8903810 https://qweenbee.tistory.com/8904668
https://qweenbee.tistory.com/8903931 https://qweenbee.tistory.com/8905654
https://qweenbee.tistory.com/8910485(??)
가시엉겅퀴 : https://qweenbee.tistory.com/8888211 https://qweenbee.tistory.com/8894279
https://qweenbee.tistory.com/8901492 https://qweenbee.tistory.com/8904672
가시엉겅퀴 새싹 : https://qweenbee.tistory.com/8901923
바늘엉겅퀴 : https://qweenbee.tistory.com/8888935 https://qweenbee.tistory.com/8894280
https://qweenbee.tistory.com/8894539 https://qweenbee.tistory.com/8894606
https://qweenbee.tistory.com/8898380 https://qweenbee.tistory.com/8901491
https://qweenbee.tistory.com/8901925 https://qweenbee.tistory.com/8912614
바늘엉겅퀴새싹: https://qweenbee.tistory.com/8901923 https://qweenbee.tistory.com/8901493
https://qweenbee.tistory.com/8904894
고려엉겅퀴 : https://qweenbee.tistory.com/8890777 https://qweenbee.tistory.com/8887010
https://qweenbee.tistory.com/8900124 https://qweenbee.tistory.com/8898317
https://qweenbee.tistory.com/8903162 https://qweenbee.tistory.com/8903327
https://qweenbee.tistory.com/8904874 https://qweenbee.tistory.com/8909363(?)
(흰)고려엉겅퀴 : https://qweenbee.tistory.com/8892848 https://qweenbee.tistory.com/8887011
https://qweenbee.tistory.com/8909420
고려엉겅퀴새싹 : https://qweenbee.tistory.com/8904432 https://qweenbee.tistory.com/8908095
정영엉겅퀴 : https://qweenbee.tistory.com/8896535 https://qweenbee.tistory.com/8903354
https://qweenbee.tistory.com/8898861 https://qweenbee.tistory.com/8903355
https://qweenbee.tistory.com/8904669 https://qweenbee.tistory.com/8903265
https://qweenbee.tistory.com/8896535
흰잎엉겅퀴: https://qweenbee.tistory.com/8913500
물엉겅퀴 : https://qweenbee.tistory.com/8892497
물엉겅퀴 새싹 : https://qweenbee.tistory.com/8902305 https://qweenbee.tistory.com/8902978
https://qweenbee.tistory.com/8893020 https://qweenbee.tistory.com/8908028
https://qweenbee.tistory.com/8908218 https://qweenbee.tistory.com/8908753
동래엉겅퀴 : https://qweenbee.tistory.com/8903357 https://qweenbee.tistory.com/8903358
큰엉겅퀴 : https://qweenbee.tistory.com/8886742 https://qweenbee.tistory.com/8888850
https://qweenbee.tistory.com/8894541 https://qweenbee.tistory.com/8900396
https://qweenbee.tistory.com/8903038 https://qweenbee.tistory.com/8903183
https://qweenbee.tistory.com/8911353
큰엉겅퀴 새싹 : https://qweenbee.tistory.com/8896204 https://qweenbee.tistory.com/8886669
https://qweenbee.tistory.com/8903038
도깨비엉겅퀴(중국) : https://qweenbee.tistory.com/8901293
https://qweenbee.tistory.com/8901343 https://qweenbee.tistory.com/8904807
반곡엉겅퀴(신칭) : https://qweenbee.tistory.com/8888770
https://qweenbee.tistory.com/8886618 https://qweenbee.tistory.com/8909363
반곡엉겅퀴 새싹 : https://qweenbee.tistory.com/8909364
반곡엉겅퀴 논문 정리 : https://qweenbee.tistory.com/8913495
지느러미엉겅퀴: https://qweenbee.tistory.com/8887126 https://qweenbee.tistory.com/8893693
https://qweenbee.tistory.com/8900982 https://qweenbee.tistory.com/8892498
https://qweenbee.tistory.com/8902608
지느러미엉겅퀴 새싹 : https://qweenbee.tistory.com/8899088
https://qweenbee.tistory.com/8900671 https://qweenbee.tistory.com/8902201
https://qweenbee.tistory.com/8906996 https://qweenbee.tistory.com/8908241
버들잎엉겅퀴 : https://qweenbee.tistory.com/8890904 https://qweenbee.tistory.com/8909509
https://qweenbee.tistory.com/m/8909562 https://qweenbee.tistory.com/8911449
https://qweenbee.tistory.com/8911450
버들잎엉겅퀴(상주): https://qweenbee.tistory.com/8889090
https://qweenbee.tistory.com/8886958 https://qweenbee.tistory.com/8892875
https://qweenbee.tistory.com/8890955 https://qweenbee.tistory.com/8889091
https://qweenbee.tistory.com/8898332 https://qweenbee.tistory.com/8903359
https://qweenbee.tistory.com/8909608 https://qweenbee.tistory.com/8911590
캐나다엉겅퀴 : https://qweenbee.tistory.com/8892773 https://qweenbee.tistory.com/8894540
서양가시엉겅퀴 : https://qweenbee.tistory.com/8896192 https://qweenbee.tistory.com/8902991
https://qweenbee.tistory.com/8907513 https://qweenbee.tistory.com/8911589(대설산)
서양가시엉겅퀴 새싹 : https://qweenbee.tistory.com/8901215
https://qweenbee.tistory.com/8902638 https://qweenbee.tistory.com/8910703
사향엉겅퀴 : https://qweenbee.tistory.com/8902872 https://qweenbee.tistory.com/8902873
https://qweenbee.tistory.com/8911183
사향엉겅퀴 새싹 : https://qweenbee.tistory.com/8904845
키무이엉겅퀴(북해도) : https://qweenbee.tistory.com/8910691
https://qweenbee.tistory.com/8911252
치시마엉겅퀴 kamtschaticum Ledeb.(대설산)https://qweenbee.tistory.com/8911601
Cirsium esculentum(몽골): https://qweenbee.tistory.com/8898733
https://qweenbee.tistory.com/8912581 https://qweenbee.tistory.com/8913064
Cirsium serratuloides : https://qweenbee.tistory.com/8912585
Olgaea속(내몽골사막지구): https://qweenbee.tistory.com/8911593
엉겅퀴속: https://qweenbee.tistory.com/8903381(부산)
https://qweenbee.tistory.com/8907514(전남)https://qweenbee.tistory.com/8903475(전남)
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