미기록종 - 서산돌나물 Sedum tricarpum Makino

 

낙동강생물자원관 박진희 박사가 발표했다고 하길래 들어가 봤더니 아직 올려져 있지 않아서 잊어버리고 있다가 이제사 확인하였다.

 

식물분류학회기 54권 4호

 

자생 미기록종 돌나물과 서산돌나물(Sedum tricarpum) 분포 보고

박진희*·김진석1·서화정2·유수창2·유정남*

국립낙동강생물자원관 다양성보전연구실 동식물연구부, 1 한반도식물다양성연구소, 2 대전대학교 생명과학과 

돌나물과의 Sedum tricarpum Makino가 한반도 서해안 지역에 자생하는 것이 본 연구에서 처음으로 확인되었다. Sedum tricarpum은 일본의 혼슈, 시코쿠, 규슈 등 주로 일본 중남부 지역에만 분포하는 일본 고유종으로 알려져 왔으나, 중국 안휘성에서도 자생하는 것이 최근 보고되었다. 한반도에서는 이 종이 2020년 충남 서해안 지역의 산지에서 처음 발견되었으며, 다년간의 관찰 및 채집을 통해 서산 팔봉산 및 태안 백화산의 해발 200-300여 미터 높이에서 각각 200여 개체가 생육하는 것으로 조사되었다. 서산에서 처음 발견되었기에 서산돌나물로 국명을 부여하였으며, 다년생 초본이고, 잎은 호생하며 엽병이 있거나 없고, 꽃이 1-1.3 cm 정도로 크며 많이 달리고, 줄기는 비교적 적게 갈라지며, 수술은 꽃잎보다 짧고 꽃밥 색이 홍자색을 띠는 점 등의 특징으로 돌나물속 내 유사종들과 구분되었다.

  

 

 

1. Leaves verticillate on sterile stems. 2. Leaves 3-verticillate on sterile stems, oblanceolate to oblong ·························································· S. sarmentosum 돌나물

  2. Leaves 4- or 5-verticillate on sterile stems, spatulate on upper part of sterile stems ····················· S. kiangnanense 잎꽃돌나물

1. Leaves alternate or opposite on sterile stems.

    3. Leaves spatulate, obovate or suborbicular, flattened.

      4. Bulblet usually present in leaf axils ······················································· S. bulbiferum 말똥비름

      4. Bulblet absent in leaf axils. 5. Leaves opposite···································· S. makinoi 둥근잎비름 

         5. Leaves alternate.

           6. Leaf apex emarginate ······························································ S. tosaense 주걱비름

           6. Leaf apex round to obtuse

             7. Plants biennial, stem trichotomously branched ······························· S. formosanum 주걱잎갯비름

             7. Plants perennial, stem irregularly branched ··································· S. tricarpum 서산돌나물

  3. Leaves linear, lanceolate, or oblong, terete or flattened.

     8. Leaves terete, apex round; petals 1.5–2 times longer than sepals.

       9. Leave linear, 8–18 mm long, somewhat sparsely arranged ····························· S. oryzifolium 땅채송화

       9. Leave widely linear-elliptic to oblong, 3–7 mm long, mostly densely arranged ·········· S. japonicum 돌채송화

    8. Leaves flattened, apex acute-acuminate; petals 2.5–4 times longer than sepals.

      10. Leaves somewhat sparsely arranged in entire stems at anthesis ·················· S. polytrichoides 바위채송화

10. Leaves densely arranged in upper half of stems at anthesis ··························· S. lepidopodum 갯돌나물

 

1. 영양줄기(sterile stem)에 잎이 돌려난다

   2. 3장의 잎이 돌려나며 긴 타원형 또는 거꿀피침모양 이다 ··········································  돌나물

   2. 4-5장의 잎이 돌려나며 줄기 상부의 잎은 주걱형 이다 ·········································· 잎꽃돌나물

1. 영양줄기(sterile stem)에 잎이 어긋나거나 마주난다

     3. 잎이 납작하며 주걱형, 달걀형 또는 타원형 이다

       4. 잎겨드랑이에 살눈이 있다 ···························································· 말똥비름

       4. 잎겨드랑이에 살눈이 없다

         5. 잎이 마주난다 ···················································································· 둥근잎비름

         5. 잎이 어긋난다

           6. 잎 끝이 오목하다다 ······································································· ·· 주걱비름

           6. 잎끝이 완만하다

             7. 두해살이풀 이며 가지는 3갈래로 난다 ···························································· 주걱잎갯비름

             7. 여러해살이풀 이며 가지는 불규칙하게 난다 ·······················································서산돌나물

   3. 잎이 선형, 창형 또는 타원형이며 원통형 이거나 납작하다

       8. 잎이 원통형이며 끝이 둥굴고 꽃잎이 꽃받침조각 보다 1.5~2배 길다

          9. 선형의 잎이 길이 8~18mm 이며 성기기게 배열된다 ············································땅채송화

          9. 피침상 선형의 잎이 길이 3~7mm 이며 촘촘하게 배열된다 ································· 돌채송화

      8. 잎이 납작하며 끝이 뾰족하고 꽃잎이 꽃받침조각 보다 2.5~4배 길다

        10. 개화기에 잎은 줄기 전체에서 다소 성기게 배열된다 ································· 바위위채송화

        10. 개화기에 잎은 줄기 전체에서 촘촘하게 배열된다 ································· 갯돌나물